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The release of lipid peroxidation products and oxidative stress signals during ferroptosis activates inflammatory pathways in synovial cells, such as the NF-κB and MAPK pathways, resulting in the release of additional inflammatory factors and creating a vicious cycle (Kumar et al., 2019

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In some embodiments, the T cell costimulatory molecule is selected from OX40, CD2, CD27, CDS, ICAM-1, LFA-1/CD11a/CD18, ICOS/CD278, 4-1BB/CD137, GITR, CD30, CD40, BAFFR, HVEM, CD7, LIGHT, NKG2C, SLAMF7, NKp80, CD160, B7-H3, and CD83, or a ligand thereof